The Contribution of Tunneling Dung Beetles to Pasture Soil Nutrition

Fig. 1. Onthophagus taurus (Schreber): (A, top) male dorsal view, (A, bottom) male lateral view; (B, top) female dorsal view, (B, bottom) female lateral view. Line = 8mm (5/16").



Image from Forage and Grazinglands article:
The Contribution of Tunneling Dung Beetles to Pasture Soil Nutrition