Fig. 3. Example of activity after four days (piedmont clay soil): (A) Dung-only treatment, (B) Onthophagus taurus treatment, (C) Onthophagus gazella treatment, (D) Onthophagus gazella treatment inverted to show multiple brood masses at bottom of pot.

Image from Forage and Grazinglands article:
The Contribution of Tunneling Dung Beetles to Pasture Soil Nutrition